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Zwickl DJ, Holder MT.  2004.  Model Parameterization, Prior Distributions and the General Time-Reversible Model in Bayesian Phylogenetics. Systematic Biology. 53:877-888.
Zakon HH, Zwickl DJ, Lu Y, Hillis DM.  2008.  Molecular evolution of communication signals in electric fish. JOURNAL OF EXPERIMENTAL BIOLOGY. 211:1814-1818. Abstract